diff --git a/src/KeyMap.cpp b/src/KeyMap.cpp
index 5bafed5a9b..f779cbbf4c 100644
--- a/src/KeyMap.cpp
+++ b/src/KeyMap.cpp
@@ -559,11 +559,11 @@ void KeyMap::listBindings(BindingList & list,
 		// a LFUN_COMMAND_PREFIX
 		if (key.prefixes) {
 			KeySequence seq = prefix;
-			seq.addkey(key.code, key.mod.first);
+			seq.addkey(key.code, key.mod.first, key.mod.second);
 			key.prefixes->listBindings(list, seq, tag);
 		} else {
 			KeySequence seq = prefix;
-			seq.addkey(key.code, key.mod.first);
+			seq.addkey(key.code, key.mod.first, key.mod.second);
 			list.push_back(Binding(key.func, seq, tag));
 		}
 	}
diff --git a/src/KeySequence.cpp b/src/KeySequence.cpp
index c0985b78b9..80cd6dae0e 100644
--- a/src/KeySequence.cpp
+++ b/src/KeySequence.cpp
@@ -147,16 +147,27 @@ docstring const KeySequence::print(outputFormat format, bool const untranslated)
 			break;
 		case BindFile:
 			KeyModifier mod = modifiers[i].first;
+			KeyModifier nmod = modifiers[i].second;
 			if (mod & ControlModifier)
 				buf += "C-";
+			else if (nmod & ControlModifier)
+				buf += "~C-";
 			if (mod & AltModifier)
 #if defined(USE_MACOSX_PACKAGING) || defined(USE_META_KEYBINDING)
 				buf += "A-";
+			else if (nmod & AltModifier)
+				buf += "~A-";
 			if (mod & MetaModifier)
 #endif
 				buf += "M-";
+#if defined(USE_MACOSX_PACKAGING) || defined(USE_META_KEYBINDING)
+			else if (nmod & MetaModifier)
+				buf += "~M-";
+#endif
 			if (mod & ShiftModifier)
 				buf += "S-";
+			else if (nmod & ShiftModifier)
+				buf += "~S-";
 
 			buf += from_utf8(sequence[i].getSymbolName());
 			break;